Publications of Kermit Ritland

Pelgas B., Bousquet J., Meirmans P., Ritland K. and Isabel N  ( 2010).  QTL mapping in white spruce: gene maps and genomic regions underlying adaptive traits related to phenology and growth across pedigrees, years, and environments, BMC Genomics (submitted)

Ritland K, Bousquet J, Isabel N, Tsumura, Krutovsky, K (2010). Genetic and comparative mapping in conifers. In Genomics of Industrial Crops: Conifers (ed. C. Kole). Science Publishers (in press).

Holliday JA, Ritland K, Aitken SN (2010). Widespread, ecologically relevant genetic markers developed from association mapping of climate-related traits in Sitka spruce (Picea sitchensis). New Phytologist 188: 501-514.

Holliday JA, Yuen M, Ritland K,  Aitken SN (2010).  Postglacial history of a widespread conifer produces inverse clines in selective neutrality. Molecular Ecology 19: 3857-3864 

Ally D, Ritland K. Otto SP. 2010. Aging in a long-lived clonal tree. PLoS Biology 8:8 e1000454.

Albouyeh R, Farzaneh N, Bohlmann J, Ritland K (2010). Multivariate analysis of digital gene expression profiles identifies a xylem signature of the vascular tissue of white spruce (Picea glauca).  Tree Genetics and Genomes. 6: 601-611.

Lo EYY, Stefanovic S, Ritland K, Dickinson TA (2010). Fine-scale comparisons of genetic variability in seed families of asexually and sexually reproducing Crataegus (Hawthorn; Rosaceae).  American Journal of Botany 97: 1014-1024,

Hamberger B, Hall D, Yuen M, Oddy C, Hamberger B, Keeling CI, Ritland C, Ritland K, Bohlmann J (2009).  Targeted isolation, sequence assembly and characterization of two white spruce (Picea glauca) BAC clones for terpenoid synthase and cytochrome P450 genes involved in conifer defence reveal insights into a conifer genome BMC Plant Biology 2009, 9:106.

Ramsay H, Rieseberg L, Ritland K (2009). Correlation of evolutionary rate with pathway position in plant terpenoid biosynthesis.  Molecular Biology and Evolution 26 : 1045-1053.

Liewlaksaneeyanawin C, Zhuang J, Tang M, Farzaneh N, Lueng G, Cullis C, Findlay S, Ritland C, Bohlmann J, Ritland K.(2009) Identification of COS markers in the Pinaceae. Tree Genetics and Genomes 5: 247-255. pdf

Lippert DN, Ralph GS, Philips M, White R, Smith D, Hardie D, Gershenzon J, Ritland K, Borchers CH Bohlmann J. (2009). Quantitative iTRAQ proteome and comparative transcriptome: Analysis of elicitor-Induced Norway Spruce (Picea abies) cells reveals elements of calcium signaling in the early conifer defense response. Proteomics 9:350-67. pdf

Albouyeh R, Ritland K. 2009. Estimating heritability of gene expression using parent-offspring regression with two-channel microarrays.  J. Heredity 100:114-118. pdf

Ally D, Ritland K, Otto S. 2008. Can clone size serve as a proxy for clone age? An exploration using microsatellite divergence in Populus tremuloides.  Molecular Ecology 17: 4897-4911. pdf

Ralph SF, Chun HJE, Kolosova N, Cooper D, Oddy C, Ritland C, Kirkpatrick R, Moore R, Barber S, Holt RA, Jones SJM, Marra MA, Douglas CJ, Ritland K, Bohlmann J. (2008) A conifer genomics resource of 200,000 spruce (Picea spp.) ESTs and 6,464 high-quality, sequence-finished full-length cDNAs for Sitka spruce (Picea sitchensis) BMC Genomics 2008, 9:484. pdf

Thompson SL, Bérubé Y, Bruneau A and Ritland K. 2008.  Three-gene identity coefficients demonstrate that clonal reproduction promotes inbreeding and spatial relatedness in yellow-cedar, Callitropsis nootkatensis.  Evolution 62: 2570-2579. pdf

O'Connell L , Ritland K, Thompson SL. 2008. Post-glacial colonization of western redcedar (Thuja plicata, Cupressaceae) revealed by microsatellite markers.  Botany 86: 194-203. pdf

Thompson S.L, Choe G, Ritland K, Whitton J. 2008. Cryptic sex within male-sterile polyploid populations of the Easter daisy, Townsendia hookeri.  International Journal of Plant Sciences169:183–193. pdf

Ukrainetz, N. K., Ritland, K. Mansfield, S. D. 2007 Identification of quantitative trait loci for wood quality and growth across eight full-sib coastal Douglas-fir families Tree Genetics and Genomics.  4: 159-170. pdf

Ukrainetz, N. K., Ritland, K. Mansfield, S. D. 2007 An AFLP linkage map for Douglas-fir based upon multiple full-sib families. Tree Genetics and Genomics. 4: 181-191. pdf

Bérubé Y, Zhuang J, Ralph S, Rungis D, Bohlmann J, and Ritland, K 2007. Characterization of EST-SSRs in loblolly pine and spruce. Tree Genetics & Genomics.  3: 251-259. pdf

Kelleher, Colin, Heesun Shin, Ian Bosdet, Chris Fjell, Rradman Chiu, Martin Krzywinski, Jennifer Wilkin, TongMing Yin, Stephen DiFazio,  Jennifer Asano, Jun Zhuang, Nicholas Putnam, Steven Jones, Marco Marra, Gerald Tuskan, Joerg Bohlmann, Brian Ellis, Kermit Ritland, Carl Douglas, Jacqueline Schein.  2007. A physical map of the highly heterozygous Populus genome: integration with the genome sequence and genetic map. The Plant Journal 50: 1063-1078. pdf

Kim S.H., Cheng K.M-T., Ritland C, Ritland K, Silversides FG. 2007. Inbreeding in Japanese Quail estimated by pedigree and microsatellite analyses.  J. Heredity 98: 378-381. pdf

Lippert D, Chowrira S, Ralph SG, Zhuang J, Aeschliman D, Ritland C, Ritland K, Bohlmann J. 2007.  Conifer defense against insects: Proteome analysis of Sitka spruce (Picea sitchensis) bark induced by mechanical wounding or feeding by white pine weevils (Pissodes strobi).  Proteomics 7: 248-270. pdf

Ally, D. and Ritland K.  2007. A case study: looking at the effects of fragmentation on genetic structure in different life history stages of old-growth mountain hemlock (Tsuga mertensiana) J. Heredity 98: 73-78. pdf

Friedmann M, Ralph SG, Aeschliman D, Zhuang J, Ritland K, Ellis BE, Bohlmann J, and CJ Douglas.  2007. Microarray gene expression profiling of the developmental transition from primary to secondary growth and constitutive defence in Sitka spruce (Picea sitchensis) apical shoots.  Journal of Experimental Botany 58: 593-614. pdf

Ralph SG, Yueh H, Friedmann M, Aeschliman D, Zeznik JA, Nelson CC, Butterfield YS, Kirkpatrick R, Liu J, Jones SJ, Marra MA, Douglas CJ, Ritland K, Bohlmann J. 2006.  Conifer defence against insects: microarray gene expression profiling of Sitka spruce (Picea sitchensis) induced by mechanical wounding or feeding by spruce budworms (Choristoneura occidentalis) or white pine weevils (Pissodes strobi) reveals large-scale changes of the host transcriptome. Plant Cell Environ 29:1545-70. pdf

Tuskan, GA, and 80 other authors including Ritland K.  2006.  The genome of black cottonwood, Populus trichocarpa (Torr. & Gray). Science 313: 1596-1604. pdf

Ralph S, Oddy C, Cooper D, Yueh H, Jancsik S, Kolosova N, Philippe RN, Aeschliman D, White R, Huber D, Ritland CE, Benoit F, Rigby T, Nantel A, Butterfield YS, Kirkpatrick R, Chun E, Liu J, Palmquist D, Wynhoven B, Stott J, Yang G, Barber S, Holt RA, Siddiqui A, Jones SJ, Marra MA, Ellis BE, Douglas CJ, Ritland K, Bohlmann J.  2006.  Genomics of hybrid poplar (Populus trichocarpax deltoides) interacting with forest tent caterpillars (Malacosoma disstria): normalized and full-length cDNA libraries, expressed sequence tags, and a cDNA microarray for the study of insect-induced defences in poplar. Molecular Ecolology 15:1275-97. pdf

Thompson, S. and Ritland K. 2006.  A novel mating system analysis for modes of self-oriented mating applied to diploid and polyploid arctic Easter daisies (Townsendia hookeri). Heredity  97: 119-126. pdf

Goodwille C, Ritland C, and Ritland K.  2006.  The genetic basis of floral traits associated with mating system evolution in Leptosiphon (Polemoniaceae): an analysis of quantitative trait loci. Evolution 60: 491–504. pdf

Beland JD, Krakowski J, Ritland C, Ritland K, El-Kassaby YA. 2005. Genetic structure and mating system of northern Arbutus menziesii populations.  Can. J. Bot. 83: 1581-1589. pdf

Wahbe TR, Ritland C, Bunnell FL, Ritland K. 2005.  Population genetic structure of tailed frogs (Ascaphus truei) in clearcut and old-growth stream habitats in south coastal British Columbia. Canadian Journal of Zoology 83: 1460-1468. pdf

Rungis D, Hamberger B, Bérubé Y, Wilkin J, Bohlmann J, Ritland, K. 2005. Efficient genetic mapping of nucleotide polymorphisms based upon DNA mismatch digestion. Molecular Breeding 16: 261-270. pdf

Wilkin JE, Shamoun SF, Ritland C, Ritland K, El-Kassaby, YA. 2005. Genetic diversity and population structure of Valdensinia heterodoxa, a potential biocontrol agent for salal in coastal British Columbia.  Can. J. Plant Pathology 27: 559-571. pdf

Ritland, K., Meagher, M.D., Edwards, D.G.W. and El-Kassaby, Y.A.  2005.  Isozyme variation and the conservation genetics of Garry oak.  Can. J. Bot 83: 1478-1487. pdf

Ritland K. 2005. Multilocus estimation of pairwise relatedness with dominant markers. Molecular Ecology 14: 3157-3165. pdf

Wilkin JE, Shamoun SF, Ritland C, Ritland K, El-Kassaby, YA. 2005. Population genetics of Gaultheria shallon in British Columbia and the Implications for management using biocontrol. Can. J. Bot. 83: 501-509. pdf

van Kleunen, M. and Ritland, K. 2005. Estimating heritabilities and genetic correlations with marker-based methods: an experimental test in Mimulus guttatus. J. Heredity 96: 368-375. pdf

Ehlting J, Mattheus N, Aeschliman DS, Li E, Hamberger B, Cullis IF, Kaneda M, Mansfield SD, Samuels AL, Ritland K, Ellis BE, Bohlmann J, Douglas CJ 2005. Global transcript profiling of primary stems from Arabidopsis thaliana identified candidate genes for missing links in lignin biosynthesis and transcriptional regulators of fiber differentiation. The Plant Journal 42: 618-640. pdf

Lippert D, Zhuang J, Ralph S, Ellis D, Gilbert M, Olafson R, Ritland K, Ellis B, Douglas C, Bohlmann J.  2005. Proteome analysis of early somatic embryogenesis in Picea glauca. Proteomics 5: 461-473. pdf

O’Connell, L., Russel, J. and Ritland, K. 2005. Post-pollination mechanisms promoting outcrossing in a self-fertile conifer.  Can. J. Bot. 83: 335-342. pdf

Hu, X-S., Goodwillie, C. and Ritland, K. 2004. Joining genetic linkage maps using a joint likelihood function. Theor. Appl. Genetics 109: 996–1004. pdf

O’Connell, L., Russel, J. and Ritland, K. 2004. Fine-scale estimation of outcrossing in western redcedar with microsatellite assay of bulked DNA. Heredity. 93: 443-449. pdf

van Kleunen M and Ritland K. 2004. Predicting evolution of floral traits associated with mating system in a natural plant population  J. Evol. Biol. 17: 1389-1399. pdf

Ritland, K. 2004. Pathways to plant population genomics.  In: Molecular Genetics and Ecology of Plant Adaptation. Proceedings of an International Workshop held December 11–13, 2002, in Vancouver, British Columbia, Canada. Edited by Q.C.B. Cronk, J. Whitton, R.H. Ree, and I.E.P. Taylor. NRC Research Press, Ottawa, Ontario. pp. 11–17.  pdf

Ritland, K and Travis, S. 2004.  Inferences involving individual coefficients of relatedness and inbreeding in natural populations of Abies. Forest Ecol. Management  197:171-180. pdf

Liewlaksaneeyanawin, C., Ritland, C., El-Kassaby, Y.A. and Ritland, K.  2004. Single-copy, species-transferable microsatellite markers developed from loblolly pine ESTs. Theor. Appl. Genetics 109:361-369. pdf

Travis, S. E., Proffitt, C.E., Ritland, K. 2004.  Population structure and inbreeding vary with successional stage in created Spartina alterniflora marshes.  Ecological Applications 14: 1189-1202. pdf

Iddrisu, M. and Ritland, K. 2004. Genetic variation, population structure and mating system in bigleaf maple (Acer macrophyllum Pursh).  Can. J. Bot. 82: 1817–1825. pdf

Ritland, K. and LeBlanc, M. 2004. Mating system of four inbreeding monkeyflower (Mimulus) species as revealed by “progeny-pair” analysis of highly informative microsatellite markers.  Plant Species Biology 19: 149-157. pdf

Rungis, D., Bérubé, Y., Zhang, J., Ralph, S., Ritland, C.E., Ellis, B.E., Douglas, C., Bohlmann, J. Ritland, K. 2004. Robust simple sequence repeat (SSR) markers for spruce (Picea spp.) from expressed sequence tags (ESTs). Theor. Appl. Genetics 109: 1283-1294. pdf

O’Connell, L. M. and K. Ritland. 2004. Somatic mutation rate at microsatellite loci in Thuja plicata. Journal of Heredity 95: 172-176. pdf

Kolosova, N., Miller, B., Ralph, S., Ellis, B.E., Douglas, C., Ritland, K., Bohlmann, J. 2004. Improved Protocol for Isolation of High-Quality mRNA from Conifers and Poplars. BioTechniques 36: 821-824. pdf

Wellman, H., Ritland, C. and Ritland, K. 2004. Genetic effects of domestication in western hemlock, Tsuga heterophylla.  Forest Genetics 10: 229-240. pdf

Liewlaksaneeyanawin, C. Ritland, K. Ritland, C. and El-Kassaby, Y. 2003.  Experimental analysis of the mating system of the white pine weevil, Pissodes strobi (Peck) (Coleoptera; Curculionideae).  Can. J. Zoology 81: 424-429.  pdf

Bérubé, Y, Ritland, CE and Ritland, K. 2003.  Isolation, characterization, and cross-species utility of microsatellites in yellow cedar, (Chamaecyparis nootkatensis).  Genome 46: 353-361. pdf

Lewis, K.G  Liewlaksaneeyanawin C, Alfaro RI, Ritland C, Ritland K, El-Kassaby YA.  2002 Sexual Reproduction in the White Pine Weevil (Pissodes strobi [Peck] [Coleoptera: Curculionidae]): Implications for Population Genetic Diversity J Hered 93: 165-169. pdf

Ritland, K.  2002. Estimation of of gene frequency and heterozygosity from pooled samples.  Molecular Ecology Notes 78. 2: 370-372. pdf

Marshall, H.D. and Ritland, K. 2002.  Genetic diversity and differentiation of Kermode bear populations. Molecular Ecology 11: 685-697. pdf

Ritland, K. 2002.  Extensions of models for the estimation of mating systems using n independent loci.  Heredity 88: 221-228. pdf

Marshall, H.D., Newton, C., and Ritland, K. 2002.  Chloroplast phylogeography and the evolution of highly polymorphic microsatellites in Lodgepole pine (Pinus contorta).  Theoretical and Applied Genetics 104: 367-378. pdf

Marshall, H.D., Newton, C. and Ritland, K.  2001. Sequence-repeat polymorphisms exhibit the signature of recombination in lodgepole pine.  Mol. Biol. Evol.  11: 2136-2138. pdf

Ritland, K., Newton, C., and Marshall, D. 2001. Inheritance and population structure of the white-phased “Kermode” black bear.  Current Biology 11: 1468-1472. pdf

O’Connell, L., F. Viard, John Russell and K. Ritland.  2001. The mating system in natural populations of western redcedar (Thuja plicata).  Canadian Journal of Botany 79:753-756. pdf

Ritland, C., Pape, T. and Ritland, K. 2001. Genetic structure of yellow-cedar (Chamaecyparis nootkatensis). Canadian Journal of Botany. 79: 822-828. pdf

Klaper, R. D. , K. Ritland, T. A. Mousseau, M. D. Hunter.  2001. Heritability of phenolics in a natural population of Quercus laevis using molecular markers.  J. Heredity 92: 421-426. pdf

Viard, F., El-Kassaby, Y. A. and Ritland, K.  2001. Diversity and genetic structure in populations of Pseudotsuga menziesii (Pinaceae) at chloroplast microsatellite loci.  Genome 44: 336-344. pdf

Lewis, K.G., Ritland, K., McLean, J.A., El-Kassaby, Y.A. Glaubitz, J., and Carlson, J.E.  2001. Randomly amplified polymorphic DNA (RAPD) reveals fine-scale genetic structure in Pissodes strobi (Coleoptera: Curculionidae).  Canadian Entomologist 133: 229-238. pdf

Ritland, K. Dupuis, L:A., Bunnell, F.L. Hung, W.L.Y. and Carlson, J.E. 2000. Phylogeography of the Tailed Frog (Ascaphus truei) in British Columbia.  Can. J. Zoology 78: 1749-1758. pdf

Ritland, K. 2000.  Marker-inferred relatedness as a tool for detecting heritability in nature.  Molecular Ecology 9: 1195-1204. pdf

Ally, D., El-Kassaby, Y.A. and Ritland, K.  2000. Genetic diversity, differentiation and mating system in mountain hemlock (Tsuga mertensiana) across British Columbia.  Forest Genetics 7; 97-108.

Lynch, M. and K. Ritland.  1999. Estimation of pairwise relatedness with molecular markers.  Genetics 152: 1753-1766. pdf

Travis, S. E., K. Ritland, T.G. Whitham and P. Keim. 1998.  A genetic linkage map of pinyon pine (Pinus edulis) based on amplified fragment length polymorphisms.  Theor. Appl. Genetics, 97: 871-880. pdf

Sun, M. and Ritland, K. 1998. Mating system parameters of yellow starthistle (Centaurea solstitialis), a successful colonizer in North America.  Heredity 80:225-232. pdf

Mousseau, T.A., K. Ritland & D.D. Heath. 1998. A novel method for estimating heritability using molecular markers.  Heredity 80:218-224. pdf

Awadalla, P. and Ritland, K. 1997. Microsatellite variation and evolution in the Mimulus guttatus species complex with contrasting mating systems. Molecular Biology and Evolution 14: 1023-1034. pdf

Lin, J.-Z. and Ritland, K. 1997. Quantitative trait loci differentiating the outbreeding Mimulus guttatus from the inbreeding Mimulus platycalyx Genetics 146: 1115-1121. pdf

Lin, J. and K. Ritland. 1996. The effects of selective genotyping on estimates of proportion of recombination between linked quantitative trait loci. Theoretical and Applied Genetics 93: 1261-1266. pdf

Ritland, K., Y.B. Fu and J.Z. Lin. 1996. Characterizing molecular quantitative variability for inbreeding depression. Plant Species Biology 11: 23-31. pdf

Fu, Y.B and K. Ritland. 1996. Marker-based inferences about epistasis for genes influencing inbreeding depression. Genetics 144: 339-348. pdf

Ritland, K. 1996. Estimators for pairwise relatedness and individual inbreeding coefficients. Genetical Research 67: 175-185. pdf

Ritland, K. 1996. A marker-based method for inferences about quantitative inheritance in natural populations. Evolution 50: 1062-1073.  pdf

Ritland, K. and C. Ritland. 1996. Inferences about quantitative inheritance based upon natural population structure in the common monkeyflower, Mimulus guttatus. Evolution 50: 1074-1082.  pdf

Lin, J. and Ritland, K. 1996. Construction of a genetic linkage map in the wild plant Mimulus using RAPD and isozyme markers. Genome 39: 63-70. pdf

Ritland, K. 1996. Inferring the genetic basis of inbreeding depression in plants. Genome 39: 1-8. pdf

El-Kassaby, Y. A. and Ritland, K. 1996. Impact of Selection and Breeding on the Genetic Diversity in Douglas-fir. Biodiversity and Conservation 5: 795-813. pdf

El-Kassaby, Y. A. and Ritland, K. 1996. Genetic variation in low elevation Douglas-fir of British Columbia and its relevance to gene conservation. Biodiversity and Conservation 5: 779-794.  pdf

Lin, J.-Z. and K. Ritland. 1995. Flower petals allow simpler and better isolation of DNA for plant RAPD analysis. Plant Molecular Biology Reporter 13: 210-213. pdf

Fu, Y.B. and K. Ritland. 1995. Marker-based inferences about the genetic basis of flower time in Mimulus guttatus. Hereditas 121: 267-272. pdf

Fenster, C.B., P. Diggle, S.C.H. Barrett, and K. Ritland. 1995. The genetics of floral development in Mimulus (Schrophulariaceae). Heredity 74: 258-266. pdf

Latta, R. and K. Ritland. 1994. Conditions favoring stable mixed mating systems with jointly evolving inbreeding depression. Journal of Theoretical Biology 170: 15-23.  pdf

Fu, Y.B. and K. Ritland. 1994. Marker-based inferences about fecundity genes contributing to inbreeding depression in Mimulus guttatus. Genome 37:1005- 1010. pdf

Latta, R. and K. Ritland. 1994. The relationship between inbreeding depression and prior inbreeding among populations of four Mimulus taxa. Evolution 48: 806-817. pdf

Fenster, C. and Ritland, K. 1994. Evidence for natural selection on mating system in Mimulus (Schrophulariaceae). International Journal of Plant Science 155: 588- 596. pdf

Fenster, C. and Ritland, K. 1994. Quantitative genetics of mating system divergence in the yellow monkeyflower species complex. Heredity 73: 422-435. pdf

Fu, Y.B. and K. Ritland. 1994. On estimating linkage of marker genes to viability genes controlling inbreeding depression. Theoretical and Applied Genetics 88: 925-932. pdf

El-Kassaby, Y.A., J. Russell, and K. Ritland. 1994. Mixed-mating in a seed orchard of western red cedar, Thuja plicata. J. Heredity 85: 227-231. pdf

Fu, Y.B. and K. Ritland. 1994. Evidence for partial dominance of viability genes in Mimulus guttatus (Schrophulariaceae). Genetics 136: 323-331. pdf

M. Johnston and Schoen, D. 1994. On the ,measurement of inbreeding depression. Evolution 48: 1735-1741 (appendix).

LeClerc-Potvin, C. and Ritland, K. 1994. Factors contributing to self-fertilization in Mimulus guttatus (Schropulariaceae): a field experiment. Amer. J. Bot. 82: 199- 205. pdf

Murawski, D.A., Fleming, T.H., Ritland, K. and J.L. Hamrick. 1994. The mating system of an autotetraploid cactus, Pachycereus pringlei. Heredity 72:86-94. pdf

Ritland, C., K. Ritland and N. Straus. 1993. Polymorphism for the ribosomal internal transcribed spacers (ITS1 and ITS2) among eight taxa of the Mimulus guttatus species complex. Molecular Biology and Evolution 10: 1273-1288. pdf

Latta, R. and K. Ritland. 1993. Models for the evolution of selfing under alternative modes of inheritance. Heredity 71: 1-10 pdf.

Ford, B., B. Ball and K. Ritland. 1993. Genetic and morphological evidence bearing on the evolution of members of Carex, Section Vesicariae (Cyperaceae) and their natural hybrids. Canadian Journal of Botany 71: 486-500. pdf

Dole, J. and K. Ritland. 1993. Inbreeding depression in two Mimulus taxa measured by multi-generational changes of the inbreeding coefficient. Evolution 47:361-373. pdf

Fenster, C. and K. Ritland. 1992. Chloroplast DNA and isozyme diversity in two Mimulus species (Scrophulariaceae) with contrasting mating systems. American Journal of Botany 79: 1440-1447. pdf

El-Kassaby, Y. and K. Ritland. 1992. Frequency-dependent male reproductive success in a polycross of Douglas-fir. Theoretical and Applied Genetics 83: 752-758. pdf

Dudash, M. and K. Ritland. 1991. Multiple paternity and self-fertilization in relation to floral age in Mimulus guttatus. American Journal of Botany 78: 1746-1753.  pdf

Ritland, K. 1991. A genetic approach to measuring pollen discounting in natural populations. American Naturalist 138: 1049-1057. pdf

Ford, B., P. Ball and K. Ritland. 1991. Allozyme diversity and genetic relationships among North American members of short-beaked taxa of Carex, Section Vesicariae (Cyperaceae). Systematic Botany 16: 116-131. pdf

Ritland, K., D. Soltis and P. Soltis. 1990. A two-locus model for the joint estimation of intergametophytic and intragametophytic selfing rates. Heredity 65: 289-296. pdf

Ritland, K. 1990. Inferences about inbreeding depression based upon changes of the inbreeding coefficient. Evolution 44: 1230-1241. pdf

Ritland, K. 1990. A series of FORTRAN computer programs for estimating plant mating systems. Journal of Heredity 81: 235-237. pdf

Hedrick, P. and K. Ritland. 1990. Gamete disequilibrium and multilocus estimation of selfing rates. Heredity 65: 343-347. pdf

Bayer, R.J., K. Ritland and B.G. Purdy. 1990. Evidence of partial apomixes in Antennaria media (Asteraceae: Inuleae) detected by the segregation of genetic markers. American Journal of Botany 77: 1078-1083. pdf

Ritland, K. 1989. Genetic differentiation, diversity and inbreeding in the mountain monkeyflower (Mimulus caespitosus) of the Washington Cascades. Canadian Journal of Botany 67: 2017-2024. pdf Pictures of the trip: http://picasaweb.google.com/kermit.ritland/1983MimulusTilingiiCollections

Ritland, C. and K. Ritland. 1989. Variation of sex allocation among eight species of the Mimulus guttatus complex. American Journal of Botany 76: 1731-1739. pdf

Ritland, K. 1989. Correlated matings in the partial selfer, Mimulus guttatus. Evolution 43: 848-859. pdf

Ritland, K. 1988. The genetic-mating structure of subdivided populations. II. Correlated mating models. Theoretical Population Biology 34: 320-346. pdf

Loaiza-Figueroa, F., K. Ritland, J. Laborde Cancino, and S.D. Tanksley. 1989. Patterns of genetic variation in the genus Capsicum (Solanaceae) in Mexico. Plant Systematics and Evolution 165: 159-188. pdf

El-Kassaby, Y.A., K. Ritland, A. Fashler and W. Devitt. 1988. The role of reproductive phenology in the mating system of a Douglas-fir seed orchard. Silvae Genetica 37: 76-82.

Ritland, K. 1987. Definition and estimation of higher-order gene fixation indices. Genetics 117: 783-793. pdf

Ritland, K. and M.T. Clegg. 1987. Evolutionary analysis of plant DNA sequences. American Naturalist 130: S74-Sl00.  pdf

Ritland, K. and F.R. Ganders. 1987. Covariation of selfing rates with parental gene fixation indices within populations of Mimulus guttatus. Evolution 4l: 760-771. pdf

Ritland, K. and F.R. Ganders. 1987. Crossability of Mimulus guttatus in relation to components of gene fixation. Evolution 4l: 772-786. pdf

El-Kassaby, Y.A. and K. Ritland. 1986. The relationship of outcrossing rate to reproductive phenology and supplemental mass pollination in a Douglas-fir seed Orchard. Silvae Genetica 35: 240-244.

El-Kassaby, Y.A. and K. Ritland. 1986. Pollen contamination in a Douglas-fir seed orchard as detected by allozyme markers. Silvae Genetica 35: 244-229. pdf

Ritland, K. 1986. Joint maximum likelihood estimation of genetic and mating structure using open-pollinated progenies. Biometrics 42: 25-43. pdf

Ritland, K. and Y.A. El-Kassaby. 1985. The nature of inbreeding in a seed orchard of Douglas-fir as shown by an efficient multilocus model. Theoretical and Applied Genetics 71: 375-384. pdf

Ritland, K. and F. Ganders. 1985. Variation in the mating system of Bidens menziesii in relation to population substructure. Heredity 55: 235-244. pdf

Ritland, K. 1985. The genetic mating structure of subdivided populations. I. Open-mating model. Theoretical Population Biology 27: 51-74. pdf

Ritland, K. and S.K. Jain. 1984. The comparative life histories of two annual Limnanthes species in a variable environment. American Naturalist 124: 656-679. pdf

Ritland, K. and S.K. Jain. 1984. A comparative study of floral and electrophoretic variation with life history variation in Limnanthes alba (Limnanathaceae). Oecologia 63: 243-251. pdf

Ritland, K. 1984. The effective proportion of self-fertilization with consanguineous matings in inbred populations. Genetics 106: 139-152. pdf

Ritland, K. 1983. The joint evolution of seed dormancy and flowering time in annual plants living in variable environments. Theoretical Population Biol. 24: 213-243. pdf

Ritland, K. and S.K. Jain. 1981. A model for the estimation of outcrossing rate and gene frequencies using n independent loci. Heredity 47: 35-52. pdf

 

Chapters in Books

Bousquet J, Isabel N, Pelgas B, Cottrell J, Rungis D, Ritland K. 2007. Spruce. In Genome Mapping and Molecular Breeding in Plants, Volume 7 Forest Trees C. Kole (Ed.) Springer-Verlag Berlin Heidelberg , pps. 93-108.

Ritland K, Ralph S, Lippert, D, Rungis D, Bohlmann J. 2006.  A new direction for conifer genomics.  In " Landscapes, Genomics and Transgenic Conifer Forests" ed. C Williams, Springer Publishers, Dordrecht Netherlands, pps. 75-84. pdf

El-Kassaby Y.A., K.Ritland, C.E. Ritland, S.N. Aitken, J.Bohlmann, and J.Krakowski. 2005.  Forest Genetics. pp. 473-486 in S.B. Watts and L. Tolland (eds.),  Forestry Handbook for British Columbia. Fifth Edition.  University of British Columbia Forestry Undergraduate Society, Vancouver, BC.

Ritland, C. and K. Ritland.  2000.  DNA fragment markers in plants.  In "Molecular methods in ecology", pp. 208-234,  ed. Allan J. Baker, Blackwell Scientific, Oxford UK. pdf

Ritland, K. 2000.  Detecting inheritance with inferred relatedness in nature. In “Adaptive genetic variation in the wild”, eds. T.A. Mousseau, B. Sinervo and J. Endler.  Oxford University Press, Oxford. Pps. 187-199.

El-Kassaby, Y.A. and Ritland, K. 1998.  Isozymes: techniques, interpretation and applications to forest genetics.  In "Forest Genetics and Tree Breeding", pp. 213-226, ed. A.K. Mandal.  CBS Publishers and Distributors, New Delhi.

Ritland, K. and J. Eckenwalder. 1991. Polymorphism, hybridization and variable evolutionary rate in molecular phylogenies. In "Plant Molecular Systematics", pps. 404-428, eds. D. Soltis, P. Soltis and J. Doyle. Chapman and Hall.

Ritland, K. and M. Clegg. 1990. Optimal DNA sequence divergence for testing phytogenetic hypotheses. In: "Molecular Evolution", UCLA Symposia on Molecular and Cellular Biology, New Series, Volume 122. pps. 289-296, eds. M. Clegg and S. O'Brian. Alan R. Liss.

Ritland, K. 1989. Marker genes and the inference of quantitative genetic parameters in the field. In: "Population Genetics, Plant Breeding and Gene Conservation", pps. 183-201, eds. A.H.D. Brown, M.T. Clegg, S.K. Jain, B.S. Weir. Sinauer Associates. pdf

Clegg, M.T., K. Ritland and G. Zurawski. 1986. Processes of chloroplast DNA evolution. In "Evolutionary processes in theory," pps. 275-294, eds. S. Karlin and E. Nevo, Academic Press.

Ritland, K. 1983. Estimation of mating systems. In "Isozymes in Plant Genetics and Breeding" pps. 289-302, eds. S. Tanksley and T. Orton, Elsevier Scientific Publications, Amsterdam. pdf